Map of Mojave desert tortoise mark-recapture and telemetry study sites

An integrated model improves inferences about survival in the Mojave desert tortoise

Despite decades of protection under the Endangered Species Act, the Mojave desert tortoise (Gopherus agassizii) continues to decline, and inconsistent historical monitoring has made it difficult to pin down exactly why. This study combines two large, previously separate datasets — nearly 4,000 mark-recapture tortoises across 35 sites (1977-2022) and almost 2,900 radio-tracked tortoises across 22 sites (1988-2022) — into a single integrated survival model. Integrating the two data sources revealed patterns that neither dataset could show on its own: males survive better than females, juveniles and subadults survive worse than adults, and survival improves with total precipitation over the prior two winters and active seasons. The integrated approach also let us tease apart “true” survival (via telemetry) from “apparent” survival (via mark-recapture alone), estimating permanent emigration rates directly for the first time at this scale. ...

February 12, 2026 · Steven J. Hromada, Brian Folt, Kevin T. Shoemaker, and coauthors

Modeling critical habitat breadth for Gopherus tortoises

The A.P.E. lab is leading a large, multi-institution collaboration investigating the climate resilience of populations of gopher tortoise (Gopherus polyphemus) and Mojave desert tortoise (Gopherus agassizii). In this project, funded by the Department of Defense (SERDP program), we are building models of tortoise vital rates (age-structured survival, fecundity) as functions of environmental and climatic covariates and simultaneously assessing for evidence of local behavioral and physiological adaptations. We are then using these models to quantify the range of environmental change that populations can withstand in situ without the need for natural selection or long-distance movement to more suitable habitats. ...

April 25, 2020 · Kevin Shoemaker