Accepted / In Press

  • Goldman, M.R., Shoemaker, K.T., Jacobson, E.K., Thomas, L., Morris, C.D., Waitland, S., Duck, C.D., Russell, D.J.F. Accepted. A Bayesian state-space model for estimating pup production in an asynchronous breeder: the grey seal (Halichoerus grypus). Royal Society Open Science.

2026

  • Sergeyev, M., Barrile, G.M., Augustine, D.J., Porensky, L.M., Duchardt, C.J., Shoemaker, K.T., Hartway, C.R., Davidson, A.D. PDOG MAPR: a decision support tool to inform management of prairie dog ecosystems. Rangeland Ecology and Management 108:1-12. https://doi.org/10.1016/j.rama.2026.04.010 [PDF]
  • Hromada, S.J., Folt, B., Shoemaker, K.T., Standen, M., Parsons, A., Allison, L.J., Dougherty, J., Drake, K.K., Esque, T.C., Walden, M.A., Freilich, J., Lovich, J.E., McLuckie, A.M., Mitchell, C.I., Vamstad, M.S., Vandergast, A.G., Hunter, E.A., Nussear, K.E., Dickson, B.G. 2026. An integrated model improves inferences about survival in the Mojave desert tortoise. Endangered Species Research 59:1-19. https://doi.org/10.3354/esr01468 [PDF]

2025

  • Shoemaker, K.T., Reich, H.E., Williams, P.J., Osterhout, M.J., Vasquez, J.P., Beckmann, J.P., Lackey, C.W., Stewart, K.M. 2025. Late season frosts and changing snowpack may exacerbate human-bear conflicts. Ecosphere. https://doi.org/10.1002/ecs2.70379 [PDF] — press coverage
  • Shoemaker, K.T., Loope, K.J. 2025. We need better ways to re-evaluate conservation policies when they’re founded on flawed research. Proceedings of the National Academy of Sciences. https://www.pnas.org/doi/10.1073/pnas.2426166122 [PDF]
  • Borokini, I.T., France, M.D., Harmon, D., Shoemaker, K.T., Weisberg, P.J., Peacock, M.M. 2025. Seed biology and regeneration niche of the threatened cold desert perennial Ivesia webberi A. Gray. Frontiers in Plant Science. https://doi.org/10.3389/fpls.2025.1568951 [PDF]
  • Nagelson, P.B., York, R.A., Shoemaker, K.T., Foster, D.E., Stephens, S.L., and Bisbing, S.M. 2025. Repeated fuel treatments fall short of fire-adapted regeneration objectives in a Sierra Nevada mixed conifer forest, USA. Ecological Applications. https://doi.org/10.1002/eap.3075

2024

  • Loope, K.J., DeSha, J.N., Aresco, M.J., Shoemaker, K.T., Hunter, E.A. 2024. Common-garden experiment reveals outbreeding depression and region-of-origin effects on reproductive success in a frequently translocated tortoise. Animal Conservation. https://doi.org/10.1111/acv.12977 [PDF]
  • Macey, S., Myers, A., Tesauro, J., Feil, A., Polanco, K., Clark, J.A., Shoemaker, K.T. 2024. Bog Turtle (Glyptemys muhlenbergii) nesting ecology and the efficacy of predator excluders in New York. Northeastern Naturalist 31(sp12), G117-G137. https://doi.org/10.1656/045.031.s1232
  • Matocq, M.D., Hunter, E.A., Murphy, P.J. and Shoemaker, K.T. 2024. Asymmetric mate preference and reproductive interference mediate climate-induced changes in mate availability in a small mammal hybrid zone. Evolution 78:1818-1830. https://doi.org/10.1093/evolut/qpae110 [PDF]
  • Loope, K.J., Akçakaya, H.R., Shoemaker, K.T. 2024. Inflated predictions from a flawed model influenced the decision to deny federal protection for the gopher tortoise. Global Ecology and Conservation. https://doi.org/10.1016/j.gecco.2024.e03089 [PDF]
  • McElroy, C.L., Windmiller, B., Berkholtz, J., Wilder, E., Welch, J., Shoemaker, K.T., Kamm, M.D. 2024. Recovery of a Blanding’s Turtle Population through Nest Protection and Headstarting. Northeastern Naturalist. https://doi.org/10.1656/045.031.s1203 [PDF]
  • Long, M.E., Stewart, K.M., Shoemaker, K.T., Reich, H., Lackey, C.W., Beckmann, J.P. 2024. Selection of den sites and chronology of denning by black bears in the eastern Sierra Nevada and western Great Basin. Ecology and Evolution. https://doi.org/10.1002/ece3.11689 [PDF]
  • Osterhout, M.J., Blum, M.E., Wakeling, B.F., Schroeder, C.A., Shoemaker, K.T., Brockman, J.C., and Stewart, K.M. 2024. Effects of Large-Scale Gold Mining on Habitat Use and Selection by American Pronghorn. Science of the Total Environment. https://doi.org/10.1016/j.scitotenv.2024.170750 [PDF]

2023

  • Walden, M.A., Loope, K.J., Hunter, E.A., Divers, S.J., Comolli, J.R., Esque, T.C., and Shoemaker, K.T. 2023. Testosterone identifies hatchling sex for Mojave desert tortoises (Gopherus agassizii). Scientific Reports. https://doi.org/10.1038/s41598-023-41677-2 [PDF]
  • Dyck, M., Shoemaker, K.T., Dennison, C., and Popescu, V. 2023. Simulated effects of roadkill and harvest on the viability of a recovering bobcat population. Journal of Wildlife Management. https://doi.org/10.1002/jwmg.22460 [PDF]
  • Gustin, M.S., Dunham-Cheatham, S.M., Choma, N., Shoemaker, K.T., and Allen, N. 2023. Determining sources of reactive mercury compounds in Reno, Nevada, USA. Frontiers in Environmental Chemistry. https://doi.org/10.3389/fenvc.2023.1202957 [PDF]
  • Goldman, M.R., Shinderman, M., Jeffress, M.R., Rodhouse, T.J., and Shoemaker, K.T. 2023. Integrating multiple sign types to improve occupancy estimation for inconspicuous species. Ecology and Evolution. https://doi.org/10.1002/ece3.10019 [PDF]
  • Dilts, T.E., Zeller, K.A., Cushman, S.A., Larrucea, E.S., Crowell, M.M., Byer, N.W., Shoemaker, K.T., Matocq, M.D. 2023. Pygmy rabbit habitat network reveals threats and opportunities for management and conservation. Landscape Ecology. https://doi.org/10.1007/s10980-023-01672-4 [PDF]
  • Crowell, M.M., LaHue, N., Heath, E., Shoemaker, K.T., and Matocq, M.D. 2023. Detection of Rabbit Hemorrhagic Disease Virus 2 in the Pygmy Rabbit (Brachylagus idahoensis) in Nevada, USA. Journal of Wildlife Diseases. https://doi.org/10.7589/JWD-D-22-00067 [PDF]
  • Dilts, T.E., Blum, M., Shoemaker, K.T., Weisberg, P.J., Stewart, K. 2023. Improved topographic ruggedness indices more accurately model fine-scale ecological patterns. Landscape Ecology. https://doi.org/10.1007/s10980-023-01646-6 [PDF]
  • Blum, M.E., Stewart, K.M., Shoemaker, K.T., Wakeling, B., Dilts, T.E., Bennett, J.R., Cox, M., Sullivan, B.W., Bleich, V. 2023. Changes in selection of resources with reproductive state in a montane ungulate. Movement Ecology 11:20. https://doi.org/10.1186/s40462-023-00378-1 [PDF]
  • Barrile, G.M., Augustine, D.J., Porensky, L.M., Duchardt, C.J., Shoemaker, K.T., Hartway, C.R., Derner, J.D., Hunter, E.A., and Davidson, A.D. 2023. A big data-model integration approach for predicting epizootics and population recovery in a keystone species. Ecological Applications. https://doi.org/10.1002/eap.2827 [PDF]
  • Blum, M.E., Stewart, K.M., Cox, M., Shoemaker, K.T., Bennett, J.R., Sullivan, B.W., Wakeling, B.F., and Bleich, V.C. 2023. Variation in diet of desert bighorn sheep around parturition: tradeoffs associated with parturition. Frontiers in Ecology and Evolution 10, 1293. https://doi.org/10.3389/fevo.2022.1071771

2022

  • Loope, K.J., Rostal, D.C., Walden, M.A., Shoemaker, K.T., and Hunter, E.A. 2022. A comparison of non-surgical methods for sexing young gopher tortoises (Gopherus polyphemus). PeerJ 10:e13599. https://doi.org/10.7717/peerj.13599 [PDF]
  • Walden, M.A., Jania, R., Kinney, M.E., Devan-Song, A., Drake, K.K., Esque, T.C., and Shoemaker, K.T. 2022. Computed tomography for measuring body fat reserves in the threatened Mojave Desert Tortoise (Gopherus agassizii). Journal of Zoo and Wildlife Medicine 54:412-423. https://doi.org/10.1638/2020-0168 [PDF]

2021

  • Reynolds, D.S., Shoemaker, K.T., von Oettingen, S., Nager, S., Veilleaux, J.P. and Moosman, P. 2021. Integrating Multiple Survey Techniques to Document a Shifting Bat Community in the Wake of White-nose Syndrome. Journal of Fish and Wildlife Management. https://doi.org/10.3996/JFWM-20-043 [PDF]
  • Byer, N.W., Dilts, T.E., Larrucea, E.S., Crowell, M.M., Shoemaker, K.T., Weisberg, P.J., Matocq, M.D. 2021. Holocene-era landscape conditions affect genetic connectivity in a sagebrush obligate species, the pygmy rabbit (Brachylagus idahoensis). Landscape Ecology. https://doi.org/10.1007/s10980-021-01328-1 [PDF]
  • Byer, N.W., Holding, M.L., Crowell, M.M., Pierson, T.W., Dilts, T.E., Shoemaker, K.T., and Matocq, M.D. 2021. Adaptive divergence despite low effective population size in a peripherally isolated population of the pygmy rabbit, Brachylagus idahoensis. Molecular Ecology. https://doi.org/10.1111/mec.16040 [PDF]
  • Fordham, D.A., Brown, S.C., Akçakaya, H.R., Brook, B.W., Haythorne, S., Manica, A., Shoemaker, K.T., Austin, J.J., Blonder, B., Rahbek, C., Nogues-Bravo, D. 2021. Process-explicit models reveal pathway to extinction for woolly mammoth using pattern-oriented validation. Ecology Letters. https://doi.org/10.1111/ele.13911 [PDF]
  • Cook, N.A., Nicolai, C.A., Shoemaker, K.T. 2021. Inferring movements and staging locations of Canvasback (Aythya valisineria) in the Pacific Flyway. Journal of Fish and Wildlife Management. https://doi.org/10.3996/JFWM-20-032 [PDF]
  • Mitchell, C.I., Shoemaker, K.T., Esque, T.C., Vandergast, A.G., Hromada, S.J., Dutcher, K.E., Heaton, J.S., and Nussear, K.E. 2021. Integrating telemetry data at several scales with spatial capture-recapture to improve density estimates for a rare and elusive reptile. Ecosphere. https://doi.org/10.1002/ecs2.3689 [PDF]
  • Hughey, L.F., Shoemaker, K.T., Stewart, K.M., McCauley, D.J., Cushman, J.H. 2021. Effects of human-altered landscapes on a reintroduced native ungulate: patterns of resource selection at the rangeland-wildland interface. Biological Conservation 257:109086. https://doi.org/10.1016/j.biocon.2021.109086 [PDF]
  • Mitchell, C.I., Friend, D.A., Phillips, L.T., Hunter, E.A., Lovich, J.E., Agha, M., Puffer, S.R., Cummings, K.L., Medica, P.A., Esque, T.C., Nussear, K.E., Shoemaker, K.T. 2021. “Unscrambling” the drivers of egg production in the Mojave desert tortoise: Climate and individual attributes predict reproductive output. Endangered Species Research 44:217-230. https://doi.org/10.3354/esr01103

2020

  • Matthews, S., Mellin, C., Pratchett, M., Shoemaker, K.T. 2020. COTSMod: A spatially explicit metacommunity model for the management of Crown-of-thorns starfish and coral recovery. Advances in Marine Biology 87:259-290. Academic Press. https://doi.org/10.1016/bs.amb.2020.09.001
  • Macey, S.K., Vaidya, P.B., Chiu, C., Clark, J.A., and Shoemaker, K.T. 2020. Evidence for Nest-Site Fidelity but not Natal Homing in Bog Turtles (Glyptemys muhlenbergii). Journal of Herpetology. https://doi.org/10.1670/19-073
  • Crowell, M.M., Matocq, M.D., Shoemaker, K.T. 2020. Ideal trapping conditions for increased trap success of pygmy rabbits (Brachylagus idahoensis) across the Great Basin. Journal of Mammalogy. https://doi.org/10.1093/jmammal/gyaa114
  • Simmons, J.B., Campbell, T., Jerde, C.L., Chandra, S., Cowan, W., Hogan, Z., Saenz, J., Shoemaker, K.T. 2020. Population dynamics of threatened Lahontan cutthroat trout in Summit Lake. Scientific Reports. https://doi.org/10.1038/s41598-020-65992-0 [PDF]
  • Heffelfinger, L.J., Stewart, K.M., Shoemaker, K.T., Darby, N.W., and Bleich, V.C. 2020. Balancing Current and Future Reproductive Investment: Variation in Resource Selection During Stages of Reproduction in a Long-Lived Herbivore. Frontiers in Ecology and Evolution. https://doi.org/10.3389/fevo.2020.00163 [PDF]
  • Hunter, E.A., Gibbs, J.P., Cayot, L.J, Tapia, W., Quinzin, M.C., Miller, J.C., Caccone, A., and Shoemaker, K.T. 2020. Seeking compromise across competing goals in conservation translocations: The case of the “extinct” Floreana Island Galapagos giant tortoise. Journal of Applied Ecology. https://doi.org/10.1111/1365-2664.13516 [PDF]

2019

  • Bleich, V.C., Blum, M.E., Holl, S.A., Shoemaker, K.T., Sustaita, D. 2019. Habitat selection by bighorn sheep in a mesic ecosystem: the San Rafael Mountains, California, USA. California Fish and Game 105:205-224. [PDF]
  • Wakeling, B.F., Van Dellen, A, Shoemaker, K.T., Gritts, M., Tull, J.C., and Stewart, K.M. 2019. The Nature of Diversity: diversity builds resiliency - in the wild and in the workplace - but it isn’t easy. The Wildlife Professional 13(6):46-49. [PDF]
  • Keehn, J.E., Shoemaker, K.T., & Feldman, C.R. 2019. Population-level effects of wind farms on a desert lizard. The Journal of Wildlife Management 83:145-157. https://doi.org/10.1002/jwmg.21565 [PDF]
  • Hall, D.B., Shoemaker, K.T., and Greger, P.D. 2019. Western red-tailed skink (Plestiodon gilberti rubricaudatus) distribution and habitat use in southern Nevada, USA. Herpetological Conservation and Biology 14:455-469. [PDF]
  • Sullivan, B.W., Nifong, R.L., Nasto, M.K., Alvarez-Clare, S., Dencker, C., Soper, F.M., Shoemaker, K.T., Ishida, F.Y., Zaragoza-Castells, J., Davidson, E.A., Cleveland, C.C. 2019. Biogeochemical recuperation of lowland tropical forest during succession. Ecology. https://doi.org/10.1002/ecy.2641 [PDF]
  • Petersky, R., Calvin, W., Shoemaker, K.T., Weisberg, P. and Harpold, A. 2019. The sensitivity of snow ephemerality to warming climate across an arid to montane vegetation gradient. Ecohydrology 12:e2060. https://doi.org/10.1002/eco.2060 [PDF]

2018

  • Shoemaker, K.T., Heffelfinger, L.J., Jackson, N.J., Blum, M.E., Wasley, T., and Stewart, K.M. 2018. A machine-learning approach for extending classical wildlife resource selection analyses. Ecology and Evolution 8:3556-3569. https://doi.org/10.1002/ece3.3936 [PDF]
  • Davidson, A.D., Hunter, E.A., Erz, J., Lightfoot, D.C., McCarthy, A.M., Mueller, J.K., and Shoemaker, K.T. 2018. Reintroducing a keystone burrowing rodent to restore an arid North American grassland: challenges and successes. Restoration Ecology 26:909-920. https://doi.org/10.1111/rec.12671 [PDF]

2017

  • Hunter, E.A., Murphy, P.J., Matocq, M.D., and Shoemaker, K.T. 2017. Differential effects of climate on survival rates drive hybrid zone dynamics and changes in species range limits. Current Biology 27:3898-3903. https://doi.org/10.1016/j.cub.2017.11.029 [PDF]
  • Davidson, A.D., Shoemaker, K.T., Weinstein, B., Costa, G., Brooks, T., Ceballos, G., Rondinini, C., Radeloff, V., and Graham, C. 2017. Geography of current and future global mammal extinction risk. PLOS ONE 12(11), e0186934. https://doi.org/10.1371/journal.pone.0186934 [PDF]
  • Che-Castaldo, C., Jenouvrier, S., Youngflesh, C., Shoemaker, K.T., Humphries, G., Lynch, H.J. 2017. Pan-Antarctic analysis reveals the importance of stochastic forcing for Adélie penguins: How noisy is too noisy for adaptive management? Nature Communications 8(1), 832. [PDF]
  • Reynolds, D.S., Shoemaker, K.T., von Oettingen, S., Najjar, S. 2017. High Rates of Winter Activity and Arousals in Two New England Bat Species: Implications for a Reduced White-nose Syndrome Impact? The Northeastern Naturalist 24(sp7), B188-B208. https://doi.org/10.1656/045.024.s720 [PDF]
  • Hileman, E.T., King, R.B., Adamski, J.M., Anton, T.G., Bailey, R.L., Baker, S.J., Johnson, B.D., Johnson, G., Kiel, B.D., Kingsbury, B.A., Kowalski, M.J., Shoemaker, K.T., Lee, Y.M., Lentini, A.M., & Campa III, H. 2017. Climatic and geographic predictors of life history variation in Eastern Massasauga (Sistrurus catenatus): A range-wide synthesis. PLoS ONE 12(2), e0172011. https://doi.org/10.1371/journal.pone.0172011 [PDF]

2016

  • Johnson, B.D., Gibbs, J.P., Bell, T.A., & Shoemaker, K.T. 2016. Manipulation of basking sites for endangered eastern massasauga rattlesnakes. The Journal of Wildlife Management 80:803-811. https://doi.org/10.1002/jwmg.21079 [PDF]
  • Johnson, B.D., Gibbs, J.P., Shoemaker, K.T., & Cohen, J.B. 2016. Demography of a Small and Isolated Population of Eastern Massasauga Rattlesnakes (Sistrurus catenatus) Threatened by Vegetative Succession. Journal of Herpetology 50:534-540. https://doi.org/10.1670/121 [PDF]
  • Ryu, H.Y., Shoemaker, K.T., Kneip, E., Pidgeon, A.M., Heglund, P.J., Bateman, B.L., … & Akçakaya, H.R. 2016. Developing population models with data from marked individuals. Biological Conservation 197:190-199. https://doi.org/10.1016/j.biocon.2016.02.031 [PDF]

2015

  • Stanton, J.C., Shoemaker, K.T., Pearson, R.G., & Akçakaya, H.R. 2015. Warning Times for Species Extinctions Due To Climate Change. Global Change Biology 21:1066-1077. https://doi.org/10.1111/gcb.12721 [PDF]
  • Shoemaker, K.T. & Akçakaya, H.R. 2015. Inferring the nature of anthropogenic threats from long-term abundance records. Conservation Biology 29:238-249. https://doi.org/10.1111/cobi.12353 [PDF]

2014

  • Gibbs, J.P., Shoemaker, K.T., Hunter, E.A., Cayot, L., & Tapia, W. 2014. Giant Tortoise Reintroduction to Española Island, Galapagos: Demographic Outcomes and Ecosystem Implications. PLoS ONE. https://doi.org/10.1371/journal.pone.0110742 [PDF]
  • Penone, C., Davidson, A.D., Shoemaker, K.T., Graham, C., Di Marco, M., Rondinini, C., Hedges, B., & Costa, G.C. 2014. Imputation of missing data in life-history traits datasets: which approach performs the best? Methods in Ecology and Evolution 5:961-970. https://doi.org/10.1111/2041-210X.12232 [PDF]
  • Fordham, D.A., Shoemaker, K.T., Schumaker, N.H., Akçakaya, H.R., Clisby, N., & Brook, B.W. 2014. How interactions between animal movement and landscape processes modify local range dynamics and extinction risk. Biology Letters 10:20140198. https://doi.org/10.1098/rsbl.2014.0198 [PDF]
  • Davidson, A.D., Friggens, M.T., Shoemaker, K.T., Hayes, C.L., Erz, J., Duran, R. 2014. Population Dynamics of Reintroduced Gunnison’s Prairie Dogs in the Southern Portion of their Range. Journal of Wildlife Management 78:429-439. https://doi.org/10.1002/jwmg.681 [PDF]
  • Shoemaker, K.T., Breisch, A.R., Jaycox, J.W., & Gibbs, J.P. 2014. Disambiguating the Minimum Viable Population Concept: Response to Reed and McCoy. Conservation Biology 28:871-873. [PDF]
  • Pearson, R.G., Stanton, J.C., Shoemaker, K.T., Aiello-Lammens, M.E., Ersts, P.J., Horning, N., Fordham, D.A., Raxworthy, C.J., Ryu, H.Y., McNees, J., & Akçakaya, H.R. 2014. Life history and spatial traits predict extinction risk due to climate change. Nature Climate Change 4:217-221. https://doi.org/10.1038/nclimate2113 [PDF]
  • Shoemaker, K.T., Lacy, R.C., Verant, M.L., Brook, B.W., Livieri, T.M., Miller, P.S., Fordham, D.A., & Akçakaya, H.R. 2014. Effects of prey metapopulation structure on the viability of black-footed ferrets in plague-impacted landscapes: a metamodelling approach. Journal of Applied Ecology 51:735-745. https://doi.org/10.1111/1365-2664.12223 [PDF]
  • Chan, H-K, Shoemaker, K.T. & Karraker, N.E. 2014. Demography of Quasipaa frogs in China reveals high vulnerability to widespread harvest pressure. Biological Conservation 170:3-9. https://doi.org/10.1016/j.biocon.2013.12.014 [PDF]

2013

  • Shoemaker, K.T., Breisch, A.R., Jaycox, J., and Gibbs, J.P. 2013. Re-examining the minimum viable population concept for long-lived species. Conservation Biology 27:542-551. https://doi.org/10.1111/cobi.12028 [PDF]
  • Shoemaker, K.T., and Gibbs, J.P. 2013. Genetic Connectivity Among Populations of the Threatened Bog Turtle (Glyptemys muhlenbergii) and the Need for a Regional Approach to Turtle Conservation. Copeia 2013:324-331. https://doi.org/10.1643/OT-12-022 [PDF]

2010

  • Shoemaker, K.T., and Gibbs, J.P. 2010. Evaluating basking-habitat deficiency in the threatened eastern massasauga rattlesnake. Journal of Wildlife Management 74:504-513. https://doi.org/10.2193/2008-519 [PDF]

2009

  • Schlaepfer, M.A., Helenbrook, W.D., Searing, K.B., and Shoemaker, K.T. 2009. Assisted colonization: evaluating contrasting management actions (and values) in the face of uncertainty. Trends in Ecology and Evolution 24:471-472. https://doi.org/10.1016/j.tree.2009.05.008 [PDF]

Book Chapters

  • Shoemaker, K.T., Johnson, G. and Prior, K.A. 2009. Habitat manipulation as a viable conservation strategy. Pp. 221-243 in Mullin, S.J. and Seigel, R.J. (eds.) Snakes: Applied Ecology and Conservation. Cornell University Press. [PDF]